GenetixBayEST. 2009
Pests · Intermediate · 15 min read

Leafhoppers on cannabis: beet leafhopper, curly top and hopperburn

Leafhoppers matter on cannabis for one reason above all: the beet leafhopper (Circulifer tenellus) spreads beet curly top virus, which was found in 81% of symptomatic hemp samples in Colorado in 2019 and has no cure. Potato leafhopper causes hopperburn but is usually a minor pest. Protect young outdoor plants with barriers, control weed hosts, monitor with yellow sticky cards and rogue infected plants; insecticides cannot stop a virus transmitted within hours.

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Short answer. Most leafhoppers on cannabis are harmless passers-by. The one that matters is the beet leafhopper (Circulifer tenellus, also Neoaliturus tenellus), which carries beet curly top virus: incurable, worst on young plants, and found in 81% of symptomatic Colorado hemp samples in 2019. The potato leafhopper (Empoasca fabae) causes hopperburn, usually minor. Protect transplants with barriers, kill the weed hosts, watch yellow sticky cards and pull infected plants. Sprays cannot outrun a virus that moves in hours.

Which leafhoppers are found on cannabis?

Leafhoppers on cannabis are small (about 3-4 mm), wedge-shaped, jumping sap-feeders of the family Cicadellidae. Over a dozen species have been found in Colorado hemp fields, most of them transients that do not breed in the crop [1]. Three are worth knowing.

Species Where Harm to cannabis Source
Beet leafhopper, Circulifer (Neoaliturus) tenellus Arid western US: Colorado West Slope, Columbia Basin, southern Idaho, central Oregon, Utah, Arizona, New Mexico, California Only known vector of beet curly top virus to hemp; feeding itself causes little injury [1][2][3][4]
Potato leafhopper, Empoasca fabae Midwest and East, migrating north each year; also Utah Hopperburn: yellow, scorched leaf tips, curling; usually low threat [16][17][18]
Ceratagallia uhleri Colorado (most common hemp leafhopper there) Breeds on hemp but causes no significant injury and transmits nothing known [1][2]
Others: Empoasca spp., Ceratagallia spp., aster leafhopper Macrosteles quadrilineatus Pacific Northwest hemp Role unknown [3]

Glossary: leafhopper, plant virus.

How do I recognise beet curly top and hopperburn?

Beet curly top in cannabis shows as pale green to yellow foliage, upward curling and puckering of leaves, twisted new growth and stunting; mid-season infection shortens internodes into a witches' broom, and badly stunted plants may die late in the season [2][4]. Severity is greater the earlier the plant is infected [4].

Hemp shows some odd variants. Plants can be "bisymptomatic": a twisted, stunted main stem with normal-looking side branches, though virus is detectable in both [1]. Some plants appear to outgrow symptoms [1][4]. Symptoms vary with variety, growth stage, temperature and virus strain [4].

Hopperburn from potato leafhopper starts as yellowing at the tips of serrated leaves, then scorching, curling and twisting; plants may be stunted [17][18]. Under UConn's description, damage can progress from first symptoms to leaf death in four to five days [18]. Light white stippling is of little consequence [16].

Differential diagnosis: curly top mimics herbicide drift, nutrient problems and broad mite damage; confirm by PCR test [4][7]. Hopperburn resembles drought, senescence or spider mite damage, so look under the leaves for the insects [16]. See How to diagnose any problem.

Leafhopper life cycle in numbers

Both important leafhoppers pass egg (inserted into leaf or stem tissue), five nymphal instars and a winged adult [2][18]. Development thresholds are 14.4 °C for beet leafhopper and 11.4 °C for potato leafhopper in the UC IPM models [8][13].

Parameter Beet leafhopper (C. tenellus) Potato leafhopper (E. fabae) Source / conditions
Lower developmental threshold 14.4 °C (58 °F) 11.4 °C (52.5 °F); 8.4 °C immatures under fluctuating temps [8] sugar beet; [13] alfalfa lab; [14]
Upper developmental threshold 35.0 °C (95 °F) 30.0 °C (86 °F) [8][13]
Egg 111.1 °C-DD 7-14 days in summer [8]; [18]
Nymphs (5 instars) 250.0 °C-DD no DD split verified [8]
Egg to adult 361.1 °C-DD (650 °F-DD) 435 °C-DD lab; 459 field (783 / 826 °F-DD) [8][13]
Pre-oviposition 13.1 d at 20 °C; 8.2 at 25; 6.4 at 28; 5.0 at 32 no value verified [9] lab
Eggs per female 105 at 20 °C; 99 at 25; 116 at 28; 197 at 32; 163 at 19-36 °C fluctuating; 300-400 under optimal conditions no lifetime value verified [9]; [3]
Eggs per female per day (derived) ~2.2 at 25 °C; ~5.0 at 32 °C no value verified from [9]: eggs ÷ (longevity − pre-oviposition)
Female longevity 58.1 d at 20 °C; 52.7 at 25; 41.9 at 28; 44.8 at 32; 96.3 fluctuating no value verified [9]
Oviposition threshold no value verified ~9.5 °C (laid at 10 °C, not 9 °C) [15]
Generations per year ~3 in Oregon and Washington 4-5 in Utah; 2+ in Connecticut [3]; [16][18]
Intrinsic rate of increase r_m no published value found measured by Hogg (1985), highest in the warmest regime; value not retrieved [14]
Sex ratio no published value found no value verified

Sources and conditions: beet leafhopper degree-days from Idaho lab and field work on sugar beet (Harries & Douglass 1948) [8]; fecundity and longevity from constant-temperature lab work (Meyerdirk & Moratorio 1987) [9]; potato leafhopper from Illinois work on alfalfa (Kouskolekas & Decker 1966) [13]. No leafhopper life table has been measured on cannabis.

How long does a generation take at each temperature?

Constant temperature Beet leafhopper egg (days) Beet leafhopper nymphs (days) Beet leafhopper egg to adult (days) Potato leafhopper egg to adult (days)
15 °C (59 °F) ~185 ~417 not viable as a linear estimate (0.6 °C above threshold) ~121
20 °C (68 °F) 20 45 64 51
25 °C (77 °F) 10.5 24 34 32
30 °C (86 °F) 7.1 16 23 23
35 °C (95 °F) 5.4 12 17.5 above upper threshold

Computed as degree-days ÷ (temperature − lower threshold) from [8] and [13]. Linear estimates are unreliable near the threshold.

For potato leafhopper under cool fluctuating regimes (8-19, 10-21 and 13-24 °C), eggs took about 35, 29 and 16 days and nymphs about 47, 39 and 23-25 days [15].

Why do temperature and humidity matter for leafhoppers?

Warmth drives beet leafhopper reproduction hard: pre-oviposition shortened from 13.1 days at 20 °C to 5.0 days at 32 °C, and eggs per female nearly doubled from 99 at 25 °C to 197 at 32 °C [9]. Generation time drops from about 64 days at 20 °C to 23 days at 30 °C [8]. Hot, dry western summers are its home climate.

Cool weather shuts beet leafhoppers down. Arizona extension notes they become less active when daily highs fall below about 60 °F (15.6 °C) and stay on their overwintering weed or crop hosts [11]. That is why disease pressure is a spring and early-summer problem in the west.

Winter moisture in the breeding grounds matters more than your field. Colorado's beet leafhoppers are largely spring migrants from New Mexico and possibly Arizona, where winter-annual mustards feed them; a wet winter there, as in 2019, means more migrants carrying virus [1].

Potato leafhopper damage is worse in warm conditions, and drought worsens the injury [18]. Hogg (1985) found intrinsic and finite rates of increase highest in the warmest regime (23-34 °C) on a per-day basis, while gross and net reproductive rates peaked in the coolest (13-24 °C) [14].

Humidity: no published humidity effect on either species was found. Leafhoppers are rarely an indoor problem, so indoor set-points (veg 24-28 °C / 60-70% RH, flower 22-26 °C / 45-55%) are not a control lever; screening and clean mothers are.

How fast can a beet leafhopper population grow?

No intrinsic rate of increase has been published for the beet leafhopper, so a true exponential projection cannot be computed. A per-generation ceiling can be, from the lab fecundity data [9] and the degree-day model [8].

Method: maximum multiplication per generation = eggs per female × fraction female × survival. Assume a 1:1 sex ratio (assumption; no published value found) and 100% survival (impossible in reality, so this is a ceiling).

Temperature Eggs per female [9] Ceiling multiplication per generation Generation length (egg to adult + pre-oviposition) Ceiling r = ln(multiplication) ÷ generation
20 °C 105 52.5x 64.5 + 13.1 ≈ 78 days ~0.051/day (doubling ~13.6 days)
25 °C 98.9 49.5x 34.1 + 8.2 ≈ 42 days ~0.092/day (doubling ~7.5 days)
28 °C 115.7 57.9x 26.6 + 6.4 ≈ 33 days ~0.123/day (doubling ~5.6 days)
32 °C 196.9 98.5x 20.5 + 5.0 ≈ 25.5 days ~0.180/day (doubling ~3.9 days)

Derived figures, not published r_m; the generation length ignores the spread of egg laying, so these ceilings run high.

The point for growers is not the local breeding rate. CSU observed only adults in hemp fields and doubts the species reproduces on the crop [1], though a CSU lab study found beet leafhoppers survived 7-day periods on hemp and laid eggs that developed [5]. Either way, your risk is set by how many virus-carrying migrants arrive from weeds and rangeland, not by how many breed in your rows.

How does beet curly top spread to cannabis?

Beet curly top reaches cannabis when a beet leafhopper that has fed on an infected plant feeds on hemp; the leafhopper is the only insect vector in North America [1][10]. The virus is persistent and circulative but does not multiply in the insect [10]. A leafhopper can acquire it within an hour of feeding and transmit after a latent period of about four hours [10].

Retention is long. EPPO cites up to 30 days [10]; Soto and Gilbertson tracked a related curtovirus in leafhoppers held on a non-host for up to 30 days [12]; Arizona extension says leafhoppers transmit for the rest of their lives [11]. The virus is not passed to offspring [11].

Weed reservoirs feed the cycle. In Colorado, prickly lettuce, dandelion and chicory most often tested positive among 41 weed species surveyed [5]. In the Pacific Northwest, kochia, Russian thistle and tumble mustard are key leafhopper hosts [3]; in Colorado, winter-annual mustards (flixweed, pepperweed) support overwintering and kochia is where summer leafhoppers are easiest to find [1].

Other routes: grafting and vegetative propagation, such as cuttings from an infected mother, transmit the virus; tools are not thought to; preliminary evidence suggests infected seed may [4]. Two strains, BCTV-Worland and BCTV-Colorado, have been found alone and mixed in Colorado hemp [6].

Scale: BCTV was detected in 81% of hemp leaf samples from 12 Colorado counties in 2019, and earlier at western sites than northern ones [5][6]. It was formally reported on Cannabis sativa in western Colorado in 2020 [7]. In the PNW it is common in the Columbia Basin, southern Idaho and central Oregon, and less so west of the Cascades [4].

Where do leafhoppers come from and how do they get in?

Beet leafhoppers fly into cannabis fields in early spring from overwintering sites on weeds [3], and in Colorado mostly arrive as long-distance migrants from the south [1]. Potato leafhoppers overwinter in the Gulf states and arrive in the north on warm winds, reaching Connecticut from June; non-reproductive adults are carried south again in autumn [18]. Utah State lists June through harvest as their season [16].

Both are flying, wind-borne adults, so distance from source habitat matters more than hygiene. Purdue advises avoiding sites next to alfalfa for potato leafhopper [17]. For beet leafhopper, weedy rangeland and field edges with kochia, Russian thistle or mustards are the sources [1][3].

Indoors, leafhoppers are rare. The realistic indoor risk is not the insect but the virus arriving in a cutting from an infected outdoor or greenhouse mother [4].

How do I scout for leafhoppers?

Scout outdoor hemp for leafhoppers with yellow sticky cards at the field edge, checked weekly and replaced as needed, plus a sweep net through the canopy [2][20]. Colorado State says sweep netting is the best way to detect beet leafhoppers in hemp [1].

Identification is the whole job. Beet leafhoppers are 3.3-3.6 mm (about 1/8 in), pale green in summer, wedge-shaped, with no head markings [2][3]. Many other leafhoppers land on the same cards, so a hand lens or microscope is needed [2][20]. Colorado's common Ceratagallia uhleri looks superficially similar but is harmless [1].

For beet leafhopper, scout the weeds as well as the crop: regular surveys of kochia patches in western Colorado show when migrations have arrived [1]. CSU suggests treating breeding areas in nearby weeds before leafhoppers move into hemp [2].

For potato leafhopper, inspect leaf undersides from June for pale, wingless nymphs that run sideways, and check scorched leaf margins and curled leaves [16][18]. Shaking plants or sweeping collects adults [18].

Action thresholds: none established for either species in hemp [2][17]. Utah State reports potato leafhopper has generally not damaged Utah hemp enough to warrant treatment [16]. For beet leafhopper, the "threshold" is effectively any viruliferous leafhopper on a young plant, which is why prevention is all that works (Grower practice).

Glossary: sticky trap, scouting.

How do I control leafhoppers on cannabis?

For beet leafhopper and curly top, control means keeping infected leafhoppers off young plants, because once a plant is infected it cannot be cured and insecticides cannot stop a transmission that takes a couple of hours [1][2]. CSU adds that beet leafhoppers are resistant to many pesticides used on sugar beet [2].

Tactic Target Evidence
Physical barriers (row covers, insect netting) over seedlings and transplants Keeps migrants off when plants are most susceptible Recommended by CSU [2]; Grower practice for timing
Weed-host control (kochia, Russian thistle, mustards, prickly lettuce) Removes breeding and virus reservoirs "Likely the most important option" per PNW, though often impractical [3]; [5]
Reflective mulch Deters leafhoppers from landing "May reduce" spread [1][4]
Rogueing infected plants Removes virus source [4]
Trap crops (sugar beet at field edges) Concentrates leafhoppers for targeted treatment Suggested by CSU [2]; untested in hemp
Clean planting stock Stops clone-borne virus [4]

Which insecticide classes are used on leafhoppers?

Insecticides used against sap-feeding hoppers in other crops come mainly from IRAC groups 4A (neonicotinoids), 4C (sulfoximines), 3A (pyrethroids and pyrethrins), 9B (pymetrozine) and 29 (flonicamid) [19]. Most are not registered on cannabis, residues fail cannabis testing, and none prevents virus transmission [1]. Follow the label and local law; no brands or rates are given here.

For potato leafhopper nymphs on leaf undersides, contact materials such as insecticidal soap and horticultural oil are grower practice in veg; we found no hemp efficacy data. Purdue notes chemical control used in other crops is not an option for hemp producers in Indiana [17]; Utah State says approved insecticides exist if populations and damage are high [16].

Rotate by mode of action if you do treat. Glossary: mode of action, contact vs systemic.

Which natural enemies attack leafhoppers?

Leafhoppers are attacked by generalist predators (green lacewings, spiders, assassin bugs, big-eyed bugs), egg parasitoid wasps of the families Mymaridae and Trichogrammatidae, and big-headed flies (Pipunculidae) [2][3]. Conservation (flowering borders, no broad-spectrum sprays) is the only practical biological tactic.

No commercial release programme or release rate has been established for leafhoppers on cannabis (no published value found). Predators eat leafhoppers but cannot stop a single viruliferous adult from infecting a plant within hours, so biological control does not manage curly top on its own.

Can I use the environment against leafhoppers?

Outdoors, timing and site are the environmental levers against beet leafhopper. Plant away from weedy rangeland and field edges with kochia, Russian thistle and mustards [1][3]. Keep transplants under cover through the spring migration, because young plants are most vulnerable and severity is greater with early infection [2][4].

Planting-date logic from other crops: in Arizona spinach and beets, delaying planting until leafhoppers become inactive (daily highs below about 60 °F / 15.6 °C) reduces curly top [11]. For a summer crop like cannabis the equivalent is protecting plants through the main spring flight, not avoiding it. Local timing varies; use your own sticky-card records (Grower practice).

For potato leafhopper, reduce plant stress, since drought worsens hopperburn [17][18], and avoid fields next to alfalfa [17].

Cultivar: there is currently no information on hemp varietal resistance to BCTV [4]. Purdue suspects cultivars bred for western states may be more stressed, and more damaged, in the Midwest [17].

Trade-off: row covers on transplants raise temperature and humidity underneath; vent or remove them once plants are established and the main flight has passed (Grower practice).

What should I not do about leafhoppers in flower?

Do not spray broad-spectrum insecticides on flowering cannabis to "stop curly top"; the virus is already in the plant, the spray cannot prevent new transmissions and residues fail testing [1].

Do not take clones from plants with curled, yellowed or twisted new growth; cuttings carry the virus [4].

Do not keep infected plants in the field to "see if they grow out of it"; they remain virus sources even when side branches look healthy [1][4].

Do not mistake hopperburn for nutrient burn and change feeding; look under the leaves first [16].

Do not count every leafhopper on a sticky card as beet leafhopper; most species in hemp are harmless [1].

What are the common myths about leafhoppers on cannabis?

  • "Spraying leafhoppers prevents curly top." Myth. Transmission takes hours and migrants keep arriving [1][10].
  • "Curly top only spreads by leafhoppers." Myth. Grafting and cuttings also transmit it; seed transmission is under investigation [4].
  • "Hopperburn is nutrient burn." Myth. Same look, different cause; find the insects [16][17].
  • "Beet leafhoppers breed in hemp fields." Contested. CSU field observations found adults only [1]; CSU lab work showed oviposition and development on hemp [5].
  • "Every leafhopper on the card is a threat." Myth. Over a dozen species occur in Colorado hemp and only beet leafhopper transmits a known hemp pathogen [1].

Common mistakes

  • Transplanting unprotected seedlings next to weedy rangeland during the spring flight.
  • Spraying after curly top symptoms appear instead of rogueing.
  • Treating sticky-card totals as beet leafhopper counts without identification.
  • Cloning from outdoor mothers in curly top regions without testing.
  • Blaming nutrients for hopperburn.

Evidence

  • Beet leafhopper as sole vector of BCTV and its prevalence in Colorado hemp: Established [1][5][6][7][10].
  • Transmission speed (1 h acquisition, 4 h latent) and retention: Established in other crops [10][11][12].
  • Degree-day models and fecundity: Established on sugar beet and alfalfa; not measured on cannabis [8][9][13].
  • Insecticides ineffective against BCTV in hemp: Supported by CSU expert assessment [1][2].
  • Reflective mulch and row covers: Supported in principle; untested in hemp [1][2][4].
  • Hopperburn on hemp: Established as a symptom; yield impact unknown [17].
  • Whether beet leafhopper reproduces on hemp in the field: Contested [1][5].

Glossary terms used here

Questions this article answers

Do leafhoppers hurt cannabis?

Most leafhoppers found on cannabis and hemp do little harm, but two matter. The beet leafhopper (Circulifer tenellus) spreads beet curly top virus, a serious, incurable disease of outdoor hemp in the western United States. The potato leafhopper (Empoasca fabae) causes hopperburn, yellow and scorched leaf tips, in the Midwest and East. Colorado's commonest hemp leafhopper, Ceratagallia uhleri, causes no significant injury.

What is beet curly top virus in cannabis?

Beet curly top virus (BCTV) is a leafhopper-spread curtovirus that stunts outdoor cannabis and hemp, yellows and mottles leaves and twists new growth into a witches' broom. The beet leafhopper is its only vector in North America. In Colorado in 2019 it was detected in 81% of symptomatic hemp leaf samples from 12 counties. Infected plants do not recover; young plants are most vulnerable.

How fast does a beet leafhopper transmit curly top?

A beet leafhopper can pick up beet curly top virus within about one hour of feeding on an infected plant, and after a latent period of about four hours it can infect new plants, according to the EPPO datasheet. It keeps the virus for weeks, up to 30 days in one study, and extension sources say for life. Because transmission is that fast, insecticides cannot prevent infection.

How long does a beet leafhopper take to develop?

The beet leafhopper needs about 361 Celsius degree-days above 14.4 C (58 F) to go from egg to adult, using the Harries and Douglass sugar beet model listed by UC IPM. That is roughly 64 days at a steady 20 C, 34 days at 25 C and 23 days at 30 C. Eggs take about 111 degree-days and the five nymphal instars about 250.

How many eggs does a beet leafhopper lay?

In laboratory work by Meyerdirk and Moratorio (1987), beet leafhopper females laid about 105 eggs at 20 C, 99 at 25 C, 116 at 28 C and 197 at 32 C, and 163 under fluctuating 19 to 36 C greenhouse conditions. Pre-oviposition fell from 13 days at 20 C to 5 days at 32 C. The Pacific Northwest handbook gives 300 to 400 eggs under optimal conditions.

What is hopperburn on cannabis?

Hopperburn on cannabis is yellowing and scorching of leaf tips, followed by curling and twisting, caused by potato leafhopper (Empoasca fabae) feeding on the undersides of leaves. Purdue reports it on Indiana hemp in 2021 and notes that no economic threshold exists. UConn says damage can go from first symptoms to dead leaves in four to five days. Confirm by finding pale, sideways-running nymphs under affected leaves.

How do I monitor leafhoppers in hemp?

Monitor leafhoppers in hemp with yellow sticky cards at field edges, checked weekly and replaced as needed, plus a sweep net, as Colorado State and Oregon State advise. Identify beet leafhopper with a hand lens: it is 3.3 to 3.6 mm, pale green to yellow and has no head markings. Other leafhoppers are common on cards, so counts are meaningless without identification.

Can insecticides stop beet curly top in cannabis?

No. Colorado State's Whitney Cranshaw states that insecticides are ineffective against beet curly top in hemp because beet leafhoppers are highly mobile and transmit the virus within a couple of hours, and Colorado State adds that beet leafhoppers are resistant to many pesticides used on sugar beet. Management relies on barriers over young plants, weed-host control, reflective mulch and removing infected plants.

Can I get curly top indoors from clones?

Yes, potentially. The beet leafhopper is an outdoor insect, so indoor rooms rarely see new infections, but the Pacific Northwest Plant Disease Handbook lists vegetative propagation, such as cuttings from an infected mother plant, as a transmission route for beet curly top virus. Do not take clones from outdoor plants with curled, yellowed new growth, and test suspect mothers by PCR.

When are leafhoppers active on outdoor hemp?

Beet leafhoppers move into crops in early spring from overwintering weeds or, in Colorado, from migrations out of New Mexico, and complete about three generations a year in Oregon and Washington. Potato leafhoppers migrate north from the Gulf states, arriving in Connecticut from June, and Utah State lists June through harvest as their time of concern. Young transplants are most at risk.

Sources

  1. Cranshaw, W. 2020. Beet leafhopper and beet curly top virus (Insects/mites that feed on hemp). Colorado State University. webdoc.agsci.colostate.edu/hempinsects/PDFs/Curly%20Top%20Be
  2. Colorado State University. Beet leafhopper (Neoaliturus tenellus) and beet curly top virus. agsci.colostate.edu/agbio/ipm-pests/beet-leafhopper-beet-cur
  3. Pacific Northwest Insect Management Handbook. Hemp: leafhoppers. pnwhandbooks.org/node/25831
  4. Frost, K., Ocamb, C.M. Hemp (Cannabis sativa) - Curly top. Pacific Northwest Plant Disease Management Handbook. pnwhandbooks.org/plantdisease/host-disease/hemp-cannabis-sat
  5. Chiginsky, J. 2021. Beet curly top virus - beet leafhopper dynamics in hemp in Colorado. M.S. thesis, Colorado State University. mountainscholar.org/items/690318d0-f830-4090-b0a5-c1e18581d7
  6. Chiginsky, J., et al. 2021. First insights into the virus and viroid communities in hemp (Cannabis sativa). Frontiers in Agronomy 3: 778433. doi.org/10.3389/fagro.2021.778433
  7. Giladi, Y., Hadad, L., Luria, N., Cranshaw, W., Lachman, O., Dombrovsky, A. 2020. First report of beet curly top virus infecting Cannabis sativa in western Colorado. Plant Disease 104(3). doi.org/10.1094/PDIS-08-19-1656-PDN
  8. UC IPM. Beet leafhopper phenology model (Harries & Douglass 1948, Ecological Monographs 18: 47-79). ipm.ucanr.edu/weather/phenology-models-description/beet-leaf
  9. Meyerdirk, D.E., Moratorio, M.S. 1987. Circulifer tenellus (Baker), the beet leafhopper: laboratory studies on fecundity and longevity. The Canadian Entomologist 119(5): 443-447. doi.org/10.4039/Ent119443-5
  10. EPPO Global Database. Beet curly top virus (BCTV00) datasheet. gd.eppo.int/taxon/BCTV00/datasheet
  11. Nischwitz, C., Olsen, M. 2011. Beet curly top disease (Curtoviruses) in spinach and table beets in Arizona (AZ1552). University of Arizona Cooperative Extension. extension.arizona.edu/sites/extension.arizona.edu/files/pubs
  12. Soto, M.J., Gilbertson, R.L. 2003. Distribution and rate of movement of the curtovirus Beet mild curly top virus in the beet leafhopper. Phytopathology 93(4): 478-484. doi.org/10.1094/PHYTO.2003.93.4.478
  13. UC IPM. Potato leafhopper phenology model (Kouskolekas & Decker 1966, Ann. Entomol. Soc. Am. 59: 292-298). ipm.ucanr.edu/PHENOLOGY/ma-potato_leafhopper.html
  14. Hogg, D.B. 1985. Potato leafhopper immature development, life tables, and population dynamics under fluctuating temperature regimes. Environmental Entomology 14(3): 349-355. doi.org/10.1093/ee/14.3.349
  15. Sher, R.B., Shields, E.J. 1991. Potato leafhopper oviposition and development under cool fluctuating temperatures. Environmental Entomology 20(4): 1113-1120. academic.oup.com/ee/article/20/4/1113/2394359
  16. Utah State University Extension. Potato leafhopper (hemp). extension.usu.edu/planthealth/ipm/notes_ag/hemp-potato-leafh
  17. Bolt, M. 2021. Potato leafhopper on hemp. Purdue Pest&Crop newsletter 2021.16. extension.entm.purdue.edu/newsletters/pestandcrop/article/po
  18. Legrand, A. 2023. Beware of hopperburn. UConn Extension. extension.uconn.edu/publication/beware-of-hopperburn-2
  19. IRAC. Mode of Action Classification (online). irac-online.org/mode-of-action/classification-online/
  20. Oregon State University Extension. Monitoring for beet leafhopper adults helps halt spread of plant virus (EM 9618). extension.oregonstate.edu/catalog/em-9618-monitoring-beet-le

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